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Ecologists regularly use animal contact networks to describe interactions underlying pathogen transmission, gene flow, and information transfer. Interactions between wildlife hosts act as transmission routes for directly transmitted pathogens and vary in ways that affect transmission efficiency. Identifying drivers of contact variation can allow both contact inference and estimation of transmission dynamics despite limited data. In desert tortoises, mating strategy, burrow use and seasonal For several species, refuges such as burrows, dens, roosts, nests are an essential resource for protection from predators and extreme environmental conditions.

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